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IB DP ESS Study Notes

2.4.4 Secondary Succession

Secondary succession is an ecological phenomenon that follows the disruption of an existing ecosystem. It involves the sequential replacement of species until the ecosystem reaches its equilibrium. This process is faster and more predictable than primary succession due to the presence of pre-existing soil and sometimes, organisms. For a better understanding of how secondary succession differs from situations where no soil is present initially, refer to primary succession.

Differences Between Primary and Secondary Succession

Primary Succession

  • Occurrence: This type of succession occurs in lifeless areas where there is no soil, such as newly formed volcanic islands or rocks.
  • Soil Formation: Soil formation is a slow process, initiated by weathering and the accumulation of organic matter from decomposing pioneer species like lichens and mosses.
  • Time Frame: Primary succession takes a longer time, often several centuries to develop a mature ecosystem.
  • Species Invasion: It begins with the invasion of pioneer species which are capable of surviving in harsh conditions, gradually leading to a more hospitable environment for subsequent species.
  • Biodiversity: The biodiversity increases gradually, with complex species invading only when the conditions become favourable.

Secondary Succession

  • Occurrence: It happens in areas where an existing community has been disrupted, but the soil remains, such as after a forest fire, hurricane, or human activities like deforestation.
  • Soil: The presence of soil, often enriched with nutrients, facilitates a quicker recovery and succession process. The soil composition basics can significantly influence the speed and nature of secondary succession.
  • Time Frame: Secondary succession is faster, with ecosystems often recovering in decades.
  • Species Invasion: Begins with species that are not specially adapted to extreme conditions, as the environment is not as harsh.
  • Biodiversity: The biodiversity can quickly reach a high level due to the existing soil and seed bank. Understanding species diversity is crucial in predicting the trajectory of secondary succession.

Stages of Secondary Succession

Stage 1: Immediate Aftermath

  • Disturbance: The initial stage is marked by a significant disturbance that disrupts the existing ecosystem, leaving the land barren but with soil.
  • Remnants: Some plants, seeds, or spores often survive the disturbance, which can accelerate the recovery process.
  • Soil: The soil, enriched with nutrients from the decay of the previous ecosystem, supports the rapid growth of plants.

Stage 2: Early Successional Stage

  • Rapid Growth: This stage is characterised by the rapid growth of grasses, herbs, and other small plants that are adapted to exploit the rich soil.
  • Animal Return: Small animals and insects begin to return, contributing to pollination and seed dispersal, and initiating the re-establishment of trophic levels.
  • Ecosystem Development: Although the ecosystem starts to regain its structure, it is still far from the original state.

Stage 3: Mid Successional Stage

  • Plant Growth: Larger plants and young trees begin to appear, overshadowing the early successional plants and often leading to their decline.
  • Animal Diversity: There is a noticeable increase in animal diversity, with the return of larger animals and increased insect diversity.
  • Competition: There is increased competition for light, space, and nutrients among plant species, leading to the dominance of more competitive species.

Stage 4: Climax Community

  • Mature Trees: The growth of mature trees signifies the establishment of a stable community. The multi-layered canopy supports a diverse understory.
  • Stability: The ecosystem reaches a state of equilibrium where species composition is relatively stable, although minor successions may continue to occur. This equilibrium is part of what ecologists term dynamic equilibrium.
  • Biodiversity: The ecosystem regains its biodiversity, though it may differ from the original state before the disturbance. Secondary succession helps illustrate how ecosystems recover and adapt, providing a practical example of impact of pollution on natural habitats.

Examples of Secondary Succession

Forest Fire

  • Initial Stage: Immediately after the fire, the landscape is charred with few survivors. However, the soil is enriched with ash and nutrients, providing a fertile ground for rapid plant growth.
  • Early Successional Stage: Grasses and small plants sprout quickly, taking advantage of the nutrient-rich soil. These plants are often those that are specifically adapted to grow rapidly following fires.
  • Mid Successional Stage: As these plants mature and reproduce, shrubs and young trees begin to grow. The returning vegetation attracts a more diverse array of animals.
  • Climax Community: Over time, a new forest emerges. The specific composition of this forest can be influenced by the severity and frequency of fires, leading to the dominance of fire-resistant species.

Abandoned Agricultural Land

  • Initial Stage: After abandonment, the land is often bare with remnants of crops and is enriched with nutrients.
  • Early Successional Stage: Weeds, grasses, and other pioneer species quickly colonise the land, aided by the existing soil fertility.
  • Mid Successional Stage: Small trees and shrubs begin to appear, increasing the complexity of the plant community and attracting a more diverse animal population.
  • Climax Community: Eventually, the land returns to a forested state, hosting a complex, stable ecosystem with high biodiversity.

Key Points to Remember

  • Soil Presence: The presence of soil is a defining characteristic of secondary succession, leading to a faster recovery of the ecosystem compared to primary succession.
  • Biodiversity: The stages of secondary succession can quickly reach a high level of biodiversity due to the existing soil and seed bank.
  • Human Influence: Human activities, such as agriculture and urban development, often initiate secondary succession by disrupting existing ecosystems.

Understanding secondary succession provides insights into the resilience and adaptability of ecosystems. It underscores nature’s capacity for regeneration and recovery, albeit often resulting in a different ecological configuration from the original state. This knowledge is pivotal for informed environmental management and conservation efforts, aiding in the prediction and facilitation of natural recovery processes following disturbances.

FAQ

The frequency of disturbances has a significant impact on secondary succession and the resulting ecosystem. Frequent disturbances can prevent an ecosystem from reaching its climax community, resulting in an environment dominated by early successional species. These ecosystems often have lower biodiversity and are characterized by species adapted to frequent disturbances. In contrast, infrequent disturbances allow ecosystems to reach and maintain their climax communities, supporting higher biodiversity and complexity. The type of disturbance also matters; certain ecosystems, like those adapted to regular fires, rely on disturbances to maintain ecological balance and biodiversity.

Animal species play a crucial role in secondary succession. Initially, small animals and insects return to the disturbed area, aiding in pollination and seed dispersal, which accelerates plant growth. As the plant community develops, it provides food and habitats, attracting a more diverse array of animals. Herbivores aid in shaping the plant community through grazing, while predators help in balancing populations. Animals also contribute to soil fertility through their waste. The return and increase of animal life signify the restoration of trophic levels, leading to a more balanced and stable ecosystem. Each animal species contributes to the complexity and functionality of the recovering ecosystem.

Secondary succession often leads to enhanced soil fertility. Following a disturbance, the initial colonisers, such as grasses and small plants, rapidly grow and die, contributing organic matter to the soil. This process of accumulation and decomposition enriches the soil with nutrients. As succession progresses, the diversity of plant species increases, each contributing different types of organic matter to the soil. The returning animal life also contributes through waste and decomposition. By the time the climax community is reached, the soil is often richer and more fertile than before the disturbance, supporting a diverse and complex ecosystem.

Yes, human intervention can play a pivotal role in accelerating and guiding secondary succession. Through actions like planting native species, removing invasive species, and managing soil quality, humans can influence the direction and pace of succession. For instance, in areas affected by deforestation, reforestation projects can reintroduce native tree species, enhancing biodiversity and ecosystem services. Controlled burns are another tool, used to manage vegetation and prevent severe wildfires, promoting a healthy, diverse ecosystem. Such interventions should be carefully planned to align with the natural progression of succession and the specific ecological context of the area.

The presence of a seed bank significantly accelerates secondary succession. A seed bank consists of viable seeds buried in the soil, which remain dormant until conditions become favourable for germination. After a disturbance, such as a forest fire, these seeds quickly sprout, leading to the rapid establishment of vegetation. This initial burst of plant growth provides food and habitat for returning animals and insects, facilitating the restoration of biodiversity. The seed bank ensures that a diverse range of plant species can immediately begin to recolonise the disturbed area, laying the foundation for ecosystem recovery.

Practice Questions

Explain the differences between primary and secondary succession, providing an example for each.

Primary and secondary succession are distinct ecological processes. Primary succession occurs in lifeless areas with no soil, such as a newly formed volcanic island. It’s a slow process involving the gradual development of soil and the establishment of life, starting with pioneer species like lichens. An example is the formation of ecosystems on volcanic rocks. In contrast, secondary succession occurs in areas where an existing ecosystem is disturbed but the soil remains, like a forest after a fire. It’s quicker as it builds upon the pre-existing soil and organisms. An example is a forest regenerating after being burnt down.

Describe the stages of secondary succession following a forest fire, and explain how biodiversity changes throughout these stages.

After a forest fire, the immediate aftermath is a charred landscape with enriched soil. This initiates the early successional stage, marked by the rapid growth of grasses and small plants, exploiting the nutrient-rich soil. Biodiversity is initially low but increases as these plants create habitats for small animals and insects. The mid-successional stage sees the emergence of shrubs and young trees, increasing plant diversity and attracting a broader array of animals. By the climax community stage, mature trees establish a multi-layered canopy, and the ecosystem reaches equilibrium with high biodiversity, although the species composition might differ from the pre-fire state.

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